So, all is working now. I don't think we did anything. My coworker installed a new identical router - it didn't make things work. He put back in the old router - it still didn't allow email sending or receiving. About 1/2 hour after that email was working.
Several days after the email problem started we were unable to get our web page from our work computers. The web page is hosted by the same company that hosts our email server. Webmail also couldn't get to our company email server except through our phones. The phones were running through the same router via wi-fi and they were able to access our website via their browsers. Very confusing.
I think someone at our email and web hosting site made a change that fixed everything but I can't say for sure. Maybe it was just the act of swapping out and then back in the router.
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